A Re-Evaluation of Reineke's Rule and Stand Density Index
Authors: Pretzsch, Hans1; Biber, Peter2
Source: Forest Science,
Volume 51, Number 4, August 2005
, pp. 304-320(17)
Abstract:
The self-thinning rule established by Reineke in 1933,
N
d-1.605 (
N,
d = number of stems and quadratic mean diameter, respectively) assumes the same allometric relationship between size and density for a wide spectrum of species under self-thinning conditions. We re-evaluate this rule based on 28 fully stocked pure stands of common beech (
Fagus sylvatica L.), Norway spruce (
Picea abies [L.] Karst.), Scots pine (
Pinus sylvestris L.), and common oak (
Quercus petraea [Mattuschka] Liebl.) in Germany that date back to the year 1870. OLS regression of the model ln(
N) =
a +
b · ln(
d) results in
b values of -1.789 for common beech, -1.664 for Norway spruce, -1.593 for Scots pine, and -1.424 for common oak. The allometric coefficient for common beech differs significantly from the other species. There is also a significant difference between the
b values of Norway spruce and common oak. Except for Scots pine, the above allometric coefficients deviate significantly (common beech) and close-to-significantly (Norway spruce, common oak) from the coefficient -1.605 postulated by Reineke in 1933. To cover the speciesspecific oscillation of mortality rates, we additionally calculate the self-thinning coefficient for each survey period. Ecological implications of the species-specific scaling exponents are stressed and consequences regarding tools for regulating stand density are discussed. FOR. SCI. 51(4):304320.
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